A325 and A490 are accepted specifications for high strength bolting applications. That includes shear, tension, and combinations of shear and tension. When bolts are subjected to shear, the faying surfaces of the two materials in contact are subjected to shear-friction. The shear-friction resistance is increased by the clamping force of the bolts. The higher the strength of the bolts, the greater the friction interface.
Further, the higher strength of the bolt, the higher shear resistance of the bolt itself.
Using the two considerations, the high strength bolts under A325 and A490 are the better options compared to normal bolt strengths.