Back then, we did use them A325 bolts interchangeably with rivets but they were only used as friction bolts, like the rivets, they didn't slip where the ordinary bolts had the hole clearances to slide in. They were initially used as a replacement for field rivets, as you see in the picture; the clip angle is riveted to the supported beam.
Those nuts do not look light but I don't remember the use of square nuts. There is a possibility that they were made that way to distinguish them. Unfortunately, you cant rely on that. If the bolt heads can be confirmed as A325 from the other side, you might be able to use the modern bearing capacity since they are not partnered with rivets in the same connection, i.e. clip to supporting beam are all bolts, clip to supported beam are all rivets.
The paint job is in remarkable shape. If it hasn't been repainted, it might be red lead.
Michael.
"Science adjusts its views based on what's observed. Faith is the denial of observation so that belief can be preserved." ~ Tim Minchin