You have only scratched the surface of my worthless information. IIRC, the T1 does have the same tension allowable as A325, but I'm home today and away from my office and can't check my references. The T-Line industry (for lattice towers anyway) has used bolts in mostly shear for many years where the connection does not have to be tight. The bolts just need to be in the hole. The A394 with shorter threads were developed to keep the threads out of the shear plane and the T1 was developed to give higher shear capacity for larger loads. When I first started (over 40 years ago) there were no T1 or T0 designations, just A394 and if you wanted more capacity you went from 5/8 to 3/4 bolts.
I am only guessing, but with the larger bolt head of the A325, the tension capacity may have more safety factor with the larger head. Also the A394 typically does not use washers while I think the A325 uses them to get a better clamping force for the torque input.
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I have been called "A storehouse of worthless information" many times.